Author: Driesch, Hans, 1867-1941
Science — Philosophy
Organism (Philosophy)
Biology — Philosophy
The Science and Philosophy of the Organism
Transcriber’s notes:
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The rare spelling typos noted in the original text have been corrected silently (e.g. invividual–>individual, hyberbola–>hyperbola) but inconsistent use of the ligature æ/ae (e.g. palæontology/palaeontology), inconsistent use of alternative spellings (e.g. learned/learnt), and occasional inconsistencies of hyphenation have been left as in the original. Minor punctuation typos have been corrected silently (e.g. index entries with missing commas). The abbreviation viz. appears in both roman and italic font.
Formatting of entries in the Table of Contents does not accurately match that of the corresponding headings in the text, particularly the heading Pt.I-B-3 which contains an extraneous α.
In Figure 12 caption, multiple ditto marks have been replaced by the relevant text for greater clarity.
THE SCIENCE AND PHILOSOPHY
OF THE ORGANISM
AGENTS | |
America | The Macmillan Company 64 & 66 Fifth Avenue, New York |
Australasia | The Oxford University Press, Melbourne |
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THE
SCIENCE AND PHILOSOPHY
OF THE ORGANISM
THE GIFFORD LECTURES DELIVERED BEFORE
THE UNIVERSITY OF ABERDEEN
IN THE YEAR 1907
BY
HANS DRIESCH, Ph.D.
HEIDELBERG
LONDON
ADAM AND CHARLES BLACK
1908
All rights reserved
PREFACE
This work is not a text-book of theoretical biology; it is a systematic presentment of those biological topics which bear upon the true philosophy of nature. The book is written in a decidedly subjective manner, and it seems to me that this is just what “Gifford Lectures” ought to be. They ought never to lose, or even try to lose, their decidedly personal character.
My appointment as Gifford Lecturer, the news of which reached me in February 1906, came just at the right moment in the progress of my theoretical studies. I had always tried to improve my previous books by adding notes or altering the arrangement; I also had left a good deal of things unpublished, and thus I often hoped that I might have occasion to arrange for a new, improved, and enlarged edition of those books. This work then is the realisation of my hopes; it is, in its way, a definitive statement of all that I have to say about the Organic.
The first volume of this work, containing the lectures for 1907—though the division into “lectures” has not been preserved—consists of Parts I. and II. of Section A, “The Chief Results of Analytical Biology.” It gives in Part I. a shortened, revised, and, as I hope, improved account of what was published in my Analytische Theorie der organischen Entwickelung (1894), Die Localisation morphogenetischer Vorgänge; ein Beweis Vitalistischen Geschehens (1899), and Die organischen Regulationen (1901), though for the professed biologist the two last-named books are by no means superseded by the new work. Part II. has never been published in any systematic form before, though there are many remarks on Systematics, Darwinism, etc., in my previous papers.
The second volume—to be published in the autumn, after the delivery of the 1908 lectures—will begin with the third and concluding part of the scientific section, which is a very carefully revised and rearranged second edition of my book, Die “Seele” als elementarer Naturfactor (1903). The greater part of this volume, however, will be devoted to the “Philosophy of the Organism,” i.e. Section B, which, in my opinion, includes the most important parts of the work.
Some apology is needed for my presuming to write in English. I was led to do so by the conviction, mistaken perhaps, that the process of translation would rob the lectures of that individual and personal character which, as I said before, seems to me so much to be desired. I wished nothing to come between me and my audience. I accordingly wrote my manuscript in English, and then submitted it to linguistic revision by such skilled aid as I was able to procure at Heidelberg. My reviser tells me that if the result of his labours leaves much to be desired, it is not to be wondered at, but that, being neither a biologist nor a philosopher, he has done his best to make me presentable to the English reader. If he has failed in his troublesome task, I know that it is not for want of care and attention, and I desire here to record my sense of indebtedness to him. He wishes to remain anonymous, but I am permitted to say that, though resident in a foreign university, he is of Scottish name and English birth.
My gratitude to my friends at Aberdeen, in particular to Professor and Mrs. J. A. Thomson, for their hospitality and great kindness towards me cannot be expressed here; they all know that they succeeded in making me feel quite at home with them.
I am very much obliged to my publishers, Messrs. A. and C. Black, for their readiness to fulfil all my wishes with respect to publication.
The lectures contained in this book were written in English by a German and delivered at a Scottish university. Almost all of the ideas discussed in it were first conceived during the author’s long residence in Southern Italy. Thus this book may be witness to the truth which, I hope, will be universally recognised in the near future—that all culture, moral and intellectual and aesthetic, is not limited by the bounds of nationality.
HANS DRIESCH.
Heidelberg, 2nd January 1908.
CONTENTS OF THE FIRST VOLUME
THE PROGRAMME
PAGE | |
On Lord Gifford’s Conception of “Science” | 1 |
Natural Sciences and “Natural Theology” | 3 |
Our Philosophical Basis | 5 |
On Certain Characteristics of Biology as a Science | 9 |
The Three Different Types of Knowledge about Nature | 13 |
General Plan of these Lectures | 15 |
General Character of the Organic Form | 19 |
SECTION A.—THE CHIEF RESULTS OF ANALYTICAL BIOLOGY
PART I.—THE INDIVIDUAL ORGANISM WITH REGARD TO
FORM AND METABOLISM
A. ELEMENTARY MORPHOGENESIS— | |
Evolutio and Epigenesis in the old Sense | 25 |
The Cell | 27 |
The Egg: its Maturation and Fertilisation | 31 |
The First Developmental Processes of Echinus | 33 |
Comparative Embryology | 44 |
The First Steps of Analytical Morphogenesis | 45 |
The Limits of Pure Description in Science | 50 |
B. EXPERIMENTAL AND THEORETICAL MORPHOGENESIS— | |
1. The Foundations of the Physiology of Development. “Evolutio” and “Epigenesis” | 52 |
The Theory of Weismann | 52 |
Experimental Morphology | 56 |
The Work of Wilhelm Roux | 58 |
The Experiments on the Egg of the Sea-urchin | 59 |
On the Intimate Structure of the Protoplasm of the Germ | 65 |
On some Specificities of Organisation in Certain Germs | 70 |
General Results of the First Period of “Entwickelungsmechanik” | 71 |
Some New Results concerning Restitutions | 74 |
2. Analytical Theory of Morphogenesis | 76 |
α. THE DISTRIBUTION OF MORPHOGENIC POTENCIES | 76 |
Prospective Value and Prospective Potency | 76 |
The Potencies of the Blastomeres | 79 |
The Potencies of Elementary Organs in General | 80 |
Explicit and Implicit Potencies: Primary and Secondary Potencies | 83 |
The Morphogenetic Function of Maturation in the Light of Recent Discoveries | 85 |
The Intimate Structure of Protoplasm: Further Remarks | 88 |
The Neutrality of the Concept of “Potency” | 89 |
β. THE “MEANS” OF MORPHOGENESIS | 89 |
β′. The Internal Elementary Means of Morphogenesis | 90 |
Some Remarks on the Importance of Surface Tension in Morphogenesis | 91 |
On Growth | 93 |
On Cell-division | 94 |
β″. The External Means of Morphogenesis | 95 |
The Discoveries of Herbst | 96 |
γ. THE FORMATIVE CAUSES OR STIMULI | 99 |
The Definition of Cause | 99 |
Some Instances of Formative and Directive Stimuli | 102 |
δ. THE MORPHOGENETIC HARMONIES | 107 |
ε. ON RESTITUTIONS | 110 |
A few Remarks on Secondary Potencies and on Secondary Morphogenetic Regulations in General | 110 |
The Stimuli of Restitutions | 113 |
3. The Problem of Morphogenetic Localisation: The Theory of the Harmonious-Equipotential system—First Proof of the Autonomy of Life | 118 |
The General Problem | 118 |
The Morphogenetic “System” | 119 |
The “Harmonious-equipotential System” | 122 |
Instances of “Harmonious-equipotential Systems” | 126 |
The Problem of the Factor E | 132 |
No Explanation offered by “Means” or “Formative Stimuli” | 132 |
No Explanation offered by a Chemical Theory of Morphogenesis | 134 |
No Machine Possible Inside the Harmonious Systems | 138 |
The Autonomy of Morphogenesis proved | 142 |
“Entelechy” | 143 |
Some General Remarks on Vitalism | 145 |
The Logic of our First Proof of Vitalism | 146 |
4. On Certain other Features of Morphogenesis Advocating its Autonomy | 150 |
Harmonious-equipotential Systems formed by Wandering Cells | 151 |
On Certain Combined Types of Morphogenetic Systems | 153 |
The “Morphaesthesia” of Noll | 157 |
Restitutions of the Second Order | 158 |
On the “Equifinality” of Restitutions | 159 |
Remarks on “Retro-Differentiation” | 163 |
C. ADAPTATION— | |
Introductory Remarks on Regulations in General | 165 |
1. Morphological Adaptation | 168 |
The Limits of the Concept of Adaptation | 168 |
Adaptations to Functional Changes from Without | 172 |
True Functional Adaptation | 176 |
Theoretical Conclusions | 179 |
2. Physiological Adaptation | 184 |
Specific Adaptedness not “Adaptation” | 186 |
Primary and Secondary Adaptations in Physiology | 188 |
On Certain Pre-requisites of Adaptations in General | 189 |
On Certain Groups of Primary Physiological Adaptations | 190 |
General Remarks on Irritability | 190 |
The Regulation of Heat Production | 193 |
Primary Regulations in the Transport of Materials and Certain Phenomena of Osmotic Pressure | 194 |
Chromatic Regulations in Algae | 197 |
Metabolic Regulations | 198 |
Immunity the only Type of a Secondary Physiological Adaptation | 204 |
No General Positive Result from this Chapter | 209 |
A few Remarks on the Limits of Regulability | 212 |
D. INHERITANCE. SECOND PROOF OF THE AUTONOMY OF LIFE— | |
The Material Continuity in Inheritance | 214 |
On Certain Theories which Seek to Compare Inheritance to Memory | 216 |
The Complex-Equipotential System and its Rôle in Inheritance | 219 |
The Second Proof of Life-Autonomy. Entelechy at the Bottom of Inheritance | 224 |
The Significance of the Material Continuity in Inheritance | 227 |
The Experimental Facts about Inheritance | 228 |
The Rôle of the Nucleus in Inheritance | 233 |
Variation and Mutation | 237 |
Conclusions from the First Main Part of these Lectures | 240 |
PART II.—SYSTEMATICS AND HISTORY
A. THE PRINCIPLES OF SYSTEMATICS— | |
Rational Systematics | 243 |
Biological Systematics | 246 |
B. THE THEORY OF DESCENT— | |
1. Generalities | 250 |
The Covert Presumption of all Theories of Descent | 253 |
The Small Value of Pure Phylogeny | 255 |
History and Systematics | 257 |
2. The Principles of Darwinism | 260 |
Natural Selection | 261 |
Fluctuating Variation the Alleged Cause of Organic Divers
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